Ian Lockwood

MUSINGS, TRIP ACCOUNTS AND IMAGES FROM SOUTH ASIA

Posts Tagged ‘Ceratophora tennentii

Sri Lanka Mountain Traverse (Part IV)

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The Knuckles Range, also known as the Dumbara Hills, as seen from the east with the South West monsoon billowing up over the edges.

The Knuckles Range, also known as the Dumbara Hills, are a relative enigmatic area that was the last point on our Sri Lankan three-mountain-range traverse. Lenny and I had been on the Riverston side in October of last year with the whole family (see blogpost) but we wanted to go back with a focus on amphibians. This time, with the goal of visiting the last of the three major mountain ranges of Sri Lanka on a single journey, we planned a route that would take us to the Knuckles on a new approach.

Leaving Nuwara Eliya on a crisp morning, we drove out on the north-eastern side of Pidurutalagala, Sri Lanka’s highest peak (2,524m). Other than a crown of cloud forest, this side of the Central Highlands is heavily cultivated and populated. The road (B332) passes through Kandapola and Walapne before descending into the Mahaweli valley through the Victoria Randenigala Rantembe Sanctuary. The Randenigala dam, controlling the outflow of Sri Lanka’s longest river is a major landmark in the dry forested landscape. W the southern Mahaweli  flood plains we passed through Mahiyanganaya on posun poya and the road was crowded with families out for dansal. The surrounding area has relatively low rainfall but enjoys the fruits of the massive Mahaweli Valley irrigation system. The landscape seems to be perpetually green and gold with paddy being grown and harvested throughout the year. Driving along the plains to the east of the Knuckles, we observed dramatic activity of the south west monsoon over the hills (see B&W)cover pictures and panorama).

Knuckles_Range_&_Monsoon_from_East_3a(B&W)(MR)(06_19)

Dramatic activity of the south west monsoon over the Knuckles range north of Mahiyanganaya

The recently filled Kalu Ganga dam on the Hettipola-Rattota road.

Looking south on the steep drive up to the Pitawala Pantana and Riverston on the Hettipola-Rattota road.

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Northern valleys of the Knuckles range as seen from the road north of Riverston.

The drive up to Riverston from Hettipola took a route around the recently flooded Kalu Ganga reservoir-Sri Lanka’s newest large hydroelectric project. The town of Pallegama, that we had driven through in October 2018, is now fully submerged. When we passed through, families were removing their last door frames, roofs and materials in anticipation of the flooding. Conditions in the exquisite dry and semi evergreen forest on the ascent were parched-the area is clearly in the rain shadow of the  Knuckles.

Once again, we enjoyed the comforts and hospitality of Sir John’s Bungalow, the small boutique hotel located just below Riverston. This was low season and, generally speaking, tourists numbers in Sri Lanka were thin as a result of the Easter Sunday bombings. But we didn’t complain about being the only guests. Nadeera Weerasinghe, the manager and super naturalist at Sir John’s, warmly welcomed us and we immediately started to plan amphibian outings.

Looking_for_Nannophrys_marmorata_Pitiwala_Pantana_1(MR)(06_19)

Lenny searching (in vain) for the rare point endemic marbled streamlined frog (Nannophrys marmorata) in the Knuckles range. Conditions were simply too dry and we could not find any sign of them on this trip.

The cloud forest at Riverston provides a rich area for amphibians and reptiles. The area receives significant of rain from the South West Monsoon and is located above 1,400 meters. Riverston is named for the peak with TV transmission towers on the Ratotta-Hettipola road (B274). The saddle or ‘gap’ where the road crosses a high point is quickly becoming a local tourist point of interest and now has several temporary shops selling carbonated drinks and hot snacks. Visitors come to walk up to the towers and a few come for the biodiversity. However, the area lacks any formal conservation management and the usual issues of solid waste, noise and carelessness are growing into alarming threats to the area’s serenity. During our full day we went with Nadeera up to Riverston to look for lizards. Lenny and I were able to find several fine examples of the endemic Leaf Nosed Lizard (Ceratophora tennentii) and Nadeera tracked down a single but rather shy and difficult to photograph Knuckles Pygmy Lizard (Caphotis dumbara).

Our search for amphibians was quite successful in spite of the dry conditions. We did look for Nannophrys marmorata but their stream habitat on Pitawala Panthana was almost completely dried up (a normal situation given that this was the direst time of the year at these natural grasslands). Further down the hill, Nadeera took us on a productive night outing in the dry evergreen forest with his naturalist friend Kais (KC). Notably we found two different female Hump Nosed Lizards (Lyriocephalus scutatus) and an array of intermediate zone frogs. During our 2nd and last night I went with Nadeera up to Riverston, where the forest is of course wetter and has different species. In about 90 minutes of looking we came across almost a dozen Pseudophilautus individuals including the green P. stuarti. At the time of writing I am still working on identifying the species from both of these walks.

Pseudophilautus_sp_green_Riverston_1a(MR)(06_19)

Most likely Pseudophilautus stuartii at Riverston, Knuckles.

On June 18th after ten days in the field, Lenny and I bade goodbye to the Knuckles and headed back to Colombo. Our final leg took us down the steep Ratotta valley into Matale and then over the low hills to Kurunegala. We knew that we would have to return to find our reticent Nannophrys marmorata that had kick started the search but we felt accomplished in all that we had seen, experienced and photographed.

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Dawn panorama above the panthana on the road to Riverston.

REFERENCES (KNUCKLES)

Amphibian Survival Alliance. Web.

Bambaradeniya Channa and S P Ekanayake. A Guide to the Biodiversity of the Knuckles Forest Range. Colombo: IUCN. 2003. Print.

De Silva, Anslem. Amphibians of Sri Lanka: A Photographic Guide to Common Frogs, Toads Caecilians. Published by author, 2009. Print.

De Silva, Anslem, Ed.  The Diversity of the Dumbara Mountains. (Lyriocephalus Special Issue). November 2005. Amphibia and Reptile Research Organization of Sri Lanka. Print.

De Silva, Anslem and Kanishka Ukuwela. A Naturalist’s Guide to the Reptiles of Sri Lanka. Colombo: Vijitha Yapa Publishing, 2017. Print.

Handunnetti, Dilrukshi. “How India’s shrub frogs crossed a bridge to Sri Lanka – and changed forever.” Mongabay. 1 May 2019. Web.

Kotagama, Sarath and Gamini Ratnavira. An Illustrated Guide to the Birds of Sri Lanka. Colombo: FOGSL, 2010. Print.

Meegaskumbura, Madhava et al. “Conservation and biogeography of threatened Amphibians of Eastern Sinharaja.” Froglog. Issue 100. January 2012. Web.

Meegaskumbura, Madhava et al. “Diversification of shrub frogs (Rhacophoridae, Pseudophilautus) in Sri Lanka-Timing and geographic context.” Molecular Phylogenetics and Evolution. 2019. Web.

Protected Planet. Sri Lanka PA Boundaries. August 2019.

Singhalage Darshani, Nadeera Weerasinghe and Gehan de Silva Wijeratne. A Naturalist’s Guide to the Flowers of Sri Lanka. Colombo: Vijitha Yapa Publications, 2018. Print.

Somaweera, Ruchira and Nilusha Somaweera. Lizards of Sri Lanka. A Colour Guide with Field Keys. Frankfurt: Edition Chimaira, 2009. Print.

Knuckles Explorations

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High peaks of the Knuckles and Meemure valley from Corbet’s Gap. (April 2007)

The Knuckles or Dumbara Kanduvetiya mountain range is a vital refuge for Sri Lankan biodiversity. Cloaked in mist, both literally and figuratively, they sit at the center of the island but are one of the least-understood natural landscapes in Sri Lanka. In 2010 UNESCO recognized the Knuckles conservation area as part of the Sri Lankan Central Highlands World Heritage Site  (UNESCO). That helped draw positive attention to the area. Located north-east of Kandy, the range is spread over about 210 square kilometers and includes a collection of rugged peaks just under 2,000 meters. I’ve been interested in the Knuckles for some time, especially since there is a strong ecological and geological affinity with the Southern Western Ghats. Last term’s school break and a family road trip gave me a chance to continue my explorations that first started in 2005.

It is difficult to move through the Knuckles area, a landscape dominated by steep escarpments, craggy peaks and isolated valleys. Dense forest makes movement difficult. I have thus far visited the two different corners on four separate family trips and I feel like we are just starting to scratch the surface of getting to know the area. When we first arrived in Sri Lanka in 2005 our family took two short visits to the Corbet’s Gap side of the Knuckles range. Now, in the last two years, we have been to the Riverston area twice. I was particularly interested to observe and document parallels in the landscape and ecology of the Knuckles with the southernmost Western Ghats. I had heard anecdotal  reports that the Agasthyamalai range, one of the richest biological zones in the Western Ghats, shares affinity with the Knuckles area. Thus, I was interested to see the pantanas and see to what extent they mirrored patterns of mid-elevation grasslands in the southern Western Ghats. These links continue to drive my ongoing interest in the Knuckles.

 

The Knuckles or Dumbara range, as seen from the summit of Sri Pada on a crisp December morning in 2013.

 

On our recent visit we took an afternoon to visit the Pitawala pantana, an area of mid elevation grasslands that is home to several rare species. Most notable is the presence of a population of the rare Kirtisinghe’s rock frog or marbled streamlined frog (Nannophrys marmorata). We found tadpoles on the stream surface but did not actually see an adult. That was disappointing but it gives us a reason to revisit the area in the next year. We were give excellent guidance by Nadeera Weerasinghe the manager of Sir John’s Bungalow, the fine accommodation that we stayed at for two nights. He also helped us identify the large Knuckles Bent Toed Geckos (Crytodactylus soba) that frequented the bungalow at night. The best shot, however was found while walking with the kids on the Riverston road at night

There are several species of reptiles and amphibians that are closely associated with the Knuckles area and are, in fact, endemic to the range. The Leaf Nosed Lizard (Ceratophora tennentii) and Knuckles Pygmy Lizard (Caphotis dumbara) were on my list and with advice from Nadeera we found several individuals on the Riverston pass. In the coming months we plan to return to learn more about this fascinating corner of Sri Lanka.

Cophotis_dumbara_at_Riverston_1a(MR)(10_18)

The endemic and rare Knuckles Pygmy Lizard (Cophotis dumbara) photographed at Riverston. November 2018

Calotes_liocephalus_jv_at_Riverston_3a(MR)(10_18)

Crestless lizard juvenile (Calotes liocephalus) at Riverston.

Riverston_pantana_1a(MR)(10_18)

Shades of the shola/grasslands mosaic? These are mid-elevation pantanas (@700-1200 m) with coarse grasses on the Riverston road.

Riverston_Rd_looking_north_pan_1a(MR)(10_18)

Morning view looking north from the Riverston road. The foreground is dominated by the pantanas- native grasslands at a mid-elevation (@700-1200 m) with coarse grasses.

knuckles range elevation 2019 (mr)

Author’s map of the Knuckles region (updated version)

Knuckles_Range_dawn_pan#1(MR)(10_16)

The broad sweep of the Knuckles range seen from the border of Wasguma National Park looking due south. October 2016.

REFERENCES & PUBLICITY

Bambaradeniya Channa and S P Ekanayake. A Guide to the Biodiversity of the Knuckles Forest Range. Colombo: IUCN. 2003. Print.

De Silva, Anslem. Amphibians of Sri Lanka: A Photographic Guide to Common Frogs, Toad Caecilians. Published by author, 2009. Print.

De Silva, Anslem, Ed.  The Diversity of the Dumbara Mountains. (Lyriocephalus Special Issue). November 2005. Amphibia and Reptile Research Organization of Sri Lanka. Print.

De Silva, Anslem and Kanishka Ukuwela. A Naturalist’s Guide to the Reptiles of Sri Lanka. Colombo: Vijitha Yapa Publishing, 2017. Print.

Ekanayake, Sarath and Channa Bambaradeniya. Trekking in the Knuckles Forest: A Trekking Guide to Alugallena, Dekinda and Nitre Cave Nature Trails. Colombo: IUCN. 2003. Print.

Lakdasun Trips. “Knuckles.” ND. Web.

Lindström, Sara.  Eskil Mattsson and S.P.Nissanka. “Forest cover change in Sri Lanka: The role of small scale farmers.” Applied Geography. May 2012. Web.

Meegaskumbura, Madhava et al.  “Amphibian Research in Sri Lanka.” Froglog. (via ResearchGate). January 2014. Web.

Somaweera, Ruchira and Nilusha Somaweera. Lizards of Sri Lanka. A Colour Guide with Field Keys. Frankfurt: Edition Chimaira, 2009. Print.

Weerawardhena, Senarathge R. and Anthony P. Russell.  “Historical land-use patterns in relation to conservation strategies for the Riverstone area, the Knuckles massif, Sri Lanka: insights gained from the recovery of anuran communities.” Taprobanica. October 2012. Web.

 

Written by ianlockwood

2019-01-12 at 9:14 pm

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